The Expert Strikes Back: Interpreting Mixture-of-Experts Language Models at Expert Level

Jeremy Herbst, Stefan Wermter, Jae Hee Lee
Proceedings of the 43rd International Conference on Machine Learning, PMLR 306:42781-42820, 2026.

Abstract

Mixture-of-Experts (MoE) architectures have become the dominant choice for scaling Large Language Models (LLMs), activating only a subset of parameters per token. While MoE architectures are primarily adopted for computational efficiency, it remains an open question whether their sparsity makes them inherently easier to interpret than dense feed-forward networks (FFNs). We compare MoE experts and dense FFNs using $k$-sparse probing and find that expert neurons are consistently less polysemantic, with the gap widening as routing becomes sparser. This suggests that sparsity pressures both individual neurons and entire experts toward monosemanticity. Leveraging this finding, we zoom out from the neuron to the expert level as a more effective unit of analysis. We validate this approach by automatically interpreting hundreds of experts. This analysis allows us to resolve the debate on specialization: experts are neither broad domain specialists (e.g., biology) nor simple token-level processors. Instead, they function as fine-grained task experts, specializing in linguistic operations or semantic tasks (e.g., closing brackets in LaTeX). Our findings suggest that MoEs are inherently interpretable at the expert level, providing a clearer path toward large-scale model interpretability. Code is available at: https://github.com/jerryy33/MoE_analysis.

Cite this Paper

BibTeX
@InProceedings{pmlr-v306-herbst26a, title = {The Expert Strikes Back: Interpreting Mixture-of-Experts Language Models at Expert Level}, author = {Herbst, Jeremy and Wermter, Stefan and Lee, Jae Hee}, booktitle = {Proceedings of the 43rd International Conference on Machine Learning}, pages = {42781--42820}, year = {2026}, editor = {Zhang, Tong and Dudik, Miroslav and Jaggi, Martin and Agarwal, Alekh and Li, Sharon and Schuurmans, Dale and Zhu, Jerry and Berkenkamp, Felix and Dong, Hanze and Bietti, Alberto}, volume = {306}, series = {Proceedings of Machine Learning Research}, month = {06--11 Jul}, publisher = {PMLR}, pdf = {https://raw.githubusercontent.com/mlresearch/v306/main/assets/herbst26a/herbst26a.pdf}, url = {https://proceedings.mlr.press/v306/herbst26a.html}, abstract = {Mixture-of-Experts (MoE) architectures have become the dominant choice for scaling Large Language Models (LLMs), activating only a subset of parameters per token. While MoE architectures are primarily adopted for computational efficiency, it remains an open question whether their sparsity makes them inherently easier to interpret than dense feed-forward networks (FFNs). We compare MoE experts and dense FFNs using $k$-sparse probing and find that expert neurons are consistently less polysemantic, with the gap widening as routing becomes sparser. This suggests that sparsity pressures both individual neurons and entire experts toward monosemanticity. Leveraging this finding, we zoom out from the neuron to the expert level as a more effective unit of analysis. We validate this approach by automatically interpreting hundreds of experts. This analysis allows us to resolve the debate on specialization: experts are neither broad domain specialists (e.g., biology) nor simple token-level processors. Instead, they function as fine-grained task experts, specializing in linguistic operations or semantic tasks (e.g., closing brackets in LaTeX). Our findings suggest that MoEs are inherently interpretable at the expert level, providing a clearer path toward large-scale model interpretability. Code is available at: https://github.com/jerryy33/MoE_analysis.} }
Endnote
%0 Conference Paper %T The Expert Strikes Back: Interpreting Mixture-of-Experts Language Models at Expert Level %A Jeremy Herbst %A Stefan Wermter %A Jae Hee Lee %B Proceedings of the 43rd International Conference on Machine Learning %C Proceedings of Machine Learning Research %D 2026 %E Tong Zhang %E Miroslav Dudik %E Martin Jaggi %E Alekh Agarwal %E Sharon Li %E Dale Schuurmans %E Jerry Zhu %E Felix Berkenkamp %E Hanze Dong %E Alberto Bietti %F pmlr-v306-herbst26a %I PMLR %P 42781--42820 %U https://proceedings.mlr.press/v306/herbst26a.html %V 306 %X Mixture-of-Experts (MoE) architectures have become the dominant choice for scaling Large Language Models (LLMs), activating only a subset of parameters per token. While MoE architectures are primarily adopted for computational efficiency, it remains an open question whether their sparsity makes them inherently easier to interpret than dense feed-forward networks (FFNs). We compare MoE experts and dense FFNs using $k$-sparse probing and find that expert neurons are consistently less polysemantic, with the gap widening as routing becomes sparser. This suggests that sparsity pressures both individual neurons and entire experts toward monosemanticity. Leveraging this finding, we zoom out from the neuron to the expert level as a more effective unit of analysis. We validate this approach by automatically interpreting hundreds of experts. This analysis allows us to resolve the debate on specialization: experts are neither broad domain specialists (e.g., biology) nor simple token-level processors. Instead, they function as fine-grained task experts, specializing in linguistic operations or semantic tasks (e.g., closing brackets in LaTeX). Our findings suggest that MoEs are inherently interpretable at the expert level, providing a clearer path toward large-scale model interpretability. Code is available at: https://github.com/jerryy33/MoE_analysis.
APA
Herbst, J., Wermter, S. & Lee, J.H.. (2026). The Expert Strikes Back: Interpreting Mixture-of-Experts Language Models at Expert Level. Proceedings of the 43rd International Conference on Machine Learning, in Proceedings of Machine Learning Research 306:42781-42820 Available from https://proceedings.mlr.press/v306/herbst26a.html.

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